It would appear that in the transition from rhamphorhynchoid to pterodactyl the fifth digit of the hindlimb was lost.
http://books.google.ca/books?id=8CKYxcylOycC&pg=PA303&lpg=PA303&dq=pterosaur+foot+toes&source=bl&ots=SopVdEuG7b&sig=pfBexLavRFjQ7OfJb1Ls7fC1PRY&hl=en&ei=p3XpS-ioKsSclgfj9qDvCg&sa=X&oi=book_result&ct=result&resnum=1&ved=0CBUQ6AEwAA#v=onepage&q=pterosaur%20foot%20toes&f=true
Page 302:
"Pterodactyloid pterosaurs however do not have a fifth digit...."
Page 303:
"The two middle toes of a pterosaur are longer than the two outer ones."
"The inner three claw impressions curve and are inclined sharply outwards and the outer one curves and is inclined slightly inwards."
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