- Pterosaur (eg. Azhdarchoidea) --->
- Primitive shorebird (eg. Graculavus) -->
- Modern shorebirds - eg. plovers, oystercatchers, sandpipers (Charadriiformes/Charadrii) and storks (Ciconiidae)
Graculavus is a prehistoric bird genus that was described by O. C. Marsh.
Graculavus is the namesake for the form taxon "Graculavidae", a plesiomorphic assembly of unrelated birds that were believed to hold a key position in avian evolution as "transitional shorebirds." Two species are known: Graculavus augustus from the Western Interior Seaway and Graculavus velox from the Atlantic Seaboard.
"However, azhdarchid footprints show that their feet were relatively small, padded and slender, and thus not well suited for wading. We argue that azhdarchids were stork- or ground hornbill-like generalists, foraging in diverse environments for small animals and carrion".